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Example sentences for: neut
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Thus, the differences in α and β for w ( d ) and w neut ( d ) must be ascribed to compensatory mutations.
In the following analyses, we denote the average fraction of viable sequences that are one mutation away from a viable sequence at distance d as x neut ( d ). Similarly, we denote as x comp ( d ) the average fraction of viable sequences that are one mutation away from a non-viable sequence at distance d .
In general, we expect w ( d ) ≈ w neut ( d ) + w comp ( d ). However, method M1 has the problem that some of the mutations counted in x comp ( d ) may actually lead back onto the network of the reference sequence rather than onto a separate neutral network.
This contribution is given by [1 - γ ( d )] w ( d ) x neut ( d ). We must then obtain the contribution of additional compensatory mutations at distance d + 1. These are given by 1 - w ( d ), the number of non-viable sequences at distance d , multiplied by the probability with which these non-viables turn into viable sequences after a single mutation, x comp ( d ). Hence, we arrive at
Figure 5shows w ( d ), w neut ( d ) and w comp ( d ), estimated using method M2, for a typical example.