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Simulation studies have shown that ML is typically more accurate and robust than MP [ 81 82 ] . Thus an ML analysis of the data was conducted by employing an heuristic search in PAUP* using TBR branch swapping on a starting tree based on neighbor joining, ACCTRAN, and STEEPEST DESCENT under the TrN+G DNA substitution model [ 83 ] . This model was selected using a Hierarchical Likelihood Ratio Test (HLRT) in Modeltest version 3.06 [ 84 ] .
Likelihood parameters for the dataset were determined by Modeltest 3.06 for the TrN+G model, and included the following: base frequencies of A = 0.2482, C = 0.207, G = 0.2466, T = 0.2982; r-matrix values of A-C = 1.0000, A-G = 3.1857, A-T = 1.0000, C-G = 1.0000, C-T = 5.1655, G-T = 1.0000; and a gamma distribution of 0.5663.