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Hence, an important conclusion of this analysis is that while interactions between adjacent trophic levels lead to predictions of higher-order dynamics during the derivation of the community model (Figure 1), the act of following the coefficients from the individual trophic-level models ( sensu [ 32 ] ) allows us to observe that trophic interactions may also appear in first-order density dependence.
The Artemisia -group sensu Bremer and Humphries [ 2 ] is not monophyletic because three segregate genera Elachanthemum, Stilnolepis , and Kaschgaria are excluded from this clade.
and Seriphidium , the Bremer and Humphries [ 2 ] morphologically based cladogram placed four small genera ( Neopallasia, Turaniphytum, Mausolea, Picrothamnus ; with a total of seven species) as closest sisters ( i.e ., segregates) of Artemisia s.s . (the Artemisia -clade sensu Bremer and Humphries), whereas Seriphidium and two small genera ( Kaschgaria, Crossostephium ; 3 species total) were placed outside the Artemisia -clade as allies to Artemisia s.s . In addition, Sphaeromeria, Filifolium, Ajaniopsis , and Stilnolepis (13 species total) were sister to this clade of eight genera that included Artemisia s.s.
An ITS phylogeny of Subtribe Leucantheminae [ 71 ] also excluded these two genera from that subtribe, which is otherwise Mediterranean in distribution, congruent with a chloroplast phylogeny based on ndh F for the entire tribe [ 64 ] . The ndh F phylogeny failed to support the subtribal classification of Tribe Anthemideae sensu Bremer and Humphries, and failed to resolve the intergeneric relationships or support the monophyly of Subtribe Artemisiinae.
Altogether these 12-13 genera comprise the Artemisia -group sensu Bremer and Humphries.
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