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2002), but the in vivo relevance of these other modifications is questionable (Mukhopadhyay et al.
A simple phylogenetic tree showing the relationships between the arsenate reductases of seven Bacteria (those that had been confirmed to possess the gene at that time) was presented as part of a study that identified the arsenic resistance genes of Thiobacillus ferrooxidans (now Acidithiobacillus ferroxidans ) [ 27 ] . Saltikov and Olson [ 28 ] used probes/primers based on the E. coli ars operon to detect ars genes in natural environments, and presented a phylogenetic analysis of these genes, however their analysis was largely confined to enteric bacteria (those that were similar enough to E. coli to be detected) and they emphasized the arsB gene rather than arsC . As part of a review of microbial arsenic transformations, we recently determined a preliminary arsC phylogeny based on 19 sequences [ 29 ] . These three studies reported phylogenies that suggested a common origin for the ars genes examined [ 27 28 29 ] . An alternative view suggested by Mukhopadhyay et al.
Gel retardation assay was performed with slight modification of the protocol used by Mukhopadhyay et al [ 28 ] . The DNA fragments to be used for gel retardation assay were endlabeled with γ 32P ATP.