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As a compromise between the practicality of high doses and the sensitivity associated with realistically low doses, we propose the ID 50 . The ID 50 can be estimated using well-established nonparametric methods like Spearman-Kärber [40] or single-parameter methods [41], and there is software available, such as a freely distributed package called ID50 developed by John Spouge (http://www.ncbi.nlm.nih.gov/CBBresearch/Spouge/Virology/, which allows an automated estimation of the ID 50 from data generated in titration experiments.