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In Artemisia s.l ., the typical limb of the Anthemideae-type ray florets are reduced to a membranous vestige, giving the impression that the small capitula are composed only of disk florets, referred to as disciform capitula (inflorescence) by Bremer and Humphries [ 2 ] . In other members of the genus, the ray flowers are absent, thus the capitulum is composed only of disk florets, i.e ., discoid.
Artemisia species, in agreement with previous studies [ 34 35 ] and not in agreement with Bremer and Humphries [ 2 ] who place it near Seriphidium sect.
Monotypic Crossostephium was treated as a segregate genus of Artemisia by Bremer and Humphries [ 2 ] . It is distinct from Artemisia by the presence of a coroniform pappus, however it was once placed near A. californica by Gray [ 45 ] and Rydberg [ 47 ] on the basis of deeply-ribbed cypselas.
Altogether these 12-13 genera comprise the Artemisia -group sensu Bremer and Humphries.
According to Bremer and Humphries [ 2 ] , the Artemisia -group is monophyletic and defined by heterogamous-disciform capitula (disk florets usually bisexual and fertile, ray florets pistillate) or homogamous-discoid capitula (disk florets usually bisexual and fertile, ray florets absent), pollen with short or no spines (associated with anemophily), and thick-walled cypselas devoid of ribs (sometimes faint ribs are present in Sphaeromeria and Crossostephium ). All of the Artemisia -group genera contain species that have been variously placed within the taxonomic boundaries of Artemisia s.l.
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